Disentangling perceptual awareness from nonconscious processing in rhesus monkeys (Macaca mulatta)

Ben-Haim, Dal Monte, Fagan, Dunham, Hassin, Chang, Santos · Proceedings of the National Academy of Sciences of the United States of America · 2021 · psychology · 10.1073/pnas.2017543118 · record Ben-Haim2021-disentangling-perceptual-awareness-from-nonc
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Where this record's text came from. faceted 2026-08 by reading the locally held text (html); every evidence string verified as an exact substring of that text before merging. Claim relations is deliberately EMPTY: this PNAS article uses numeric citations throughout, so although it positions itself against prior work ('To date, no one has explored if similar performance dissociations exist in a nonhuman species.'), no prior study is named in the article text and recording a target would require importing knowledge from outside the paper (rules 2 and 7). Replication status = replicated on the strength of the paper's own statement that Experiments 3 and 4 replicate Experiments 1 and 2 in a different paradigm; this is internal replication, not external. Overall claim is also anchored by: 'Quotation withheld' 2026-08-12 and 2026-08-15 schema-0.3 passes: every quote verified as an exact substring of the staged text and anchored at merge time; concept terms verified present; every edge endpoint an existing node.
4 quotations on this page contain a reference number where the paper printed a superscript. This journal style prints citations as raised numerals — parameters12,13 — and the extraction flattened them into the line, so the sentence reads parameters 12 , 13 . The digits are citation markers and not part of what the author wrote. The quotes keep them for the same reason they keep every other extraction fault: a quote is reproduced exactly or it proves nothing.
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Other fields define these differently the same word, defined another way in another discipline
statistical-significance 2 fields · theory of mind 2 fields

Tags on this paper

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double dissociation single dissociations subliminal supraliminal nonconscious processing visual awareness spatial-cueing paradigm explanatory gap theory of mind Subliminal cues capture gaze toward the cued (wrong) side and, without awareness, the subject cannot disengage before target onset, so the eyes are still at the wrong location when the target appears and the response is slowed Conscious access to the cue lets the subject deploy a deliberate strategy that redirects gaze to the opposite location before the target appears, converting the same cue from interference into facilitation Because the two conditions push performance in opposite directions, the cross-over pattern itself is what separates aware from unaware processing in a nonverbal subject a subliminal cue cannot be disengaged from, so it drags attention the wrong way; a supraliminal one can be reasoned about, so it helps - the two pull in opposite directions the design's force comes from predicting the REVERSE of what salience alone would predict, so a confound cannot produce the result only two processing modes can produce two opposite signatures, so the crossing itself is the inference - not either half of it rhesus monkeys forced-guessing paradigm cross-over double dissociation supraliminal stimuli subliminal stimuli response time gaze frequency linear mixed model linear mixed models F-test correlation binomial test individuals’ binomial test’s P values chi-squared test standard error of the mean SEM of monkey trials mean and standard deviation learning-slope correlation anova statistical-significance binomial-test behavioural-experiment

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conceptconceptsan idea the paper works with, quoted where the paper uses it
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mechanismmechanismshow the reported effect is said to come about
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object of studyobject_of_studywhat the paper is about — the thing it investigates
phenomenonphenomenonthe effect or regularity under study
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transfertransfersa method or concept carried in from another field

The paper

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alternatives 2 · claim_relations 3 · claim_type 1 · concept_edges 4 · concepts 9 · content_tags 41 · contribution 1 · controls 1 · limitations 6 · mechanisms 6 · method 1 · object_of_study 1 · phenomenon 1 · research_question 1 · senses 4 · transfers 2

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facettagfieldconf.whose words
object_of_studyWhether rhesus monkeys show the opposite performance signatures that in humans distinguish conscious from nonconscious processing.
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object_of_studythe paper's words
research_questionSince a nonverbal animal cannot report what it is aware of, is there any behavioural test that can tell conscious from nonconscious processing in one?
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research_questionthe paper's words
contributionTurns a human double dissociation into a nonverbal test of awareness, and finds the same opposite signatures in monkeys - which is evidence for two distinct processing modes without requiring a report.
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contributionthe paper's words
claim_typeexistence
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claim_typethe paper's words
phenomenondouble dissociation of visual awareness in nonhuman primates
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phenomenonthe paper's words
methodbehavioural-experiment
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methodthe paper's words
mechanismsmechanisms
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mechanisms[0]the tagger's words
mechanismsmechanisms
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mechanisms[1]the tagger's words
mechanismsmechanisms
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mechanisms[2]the tagger's words
mechanismsmechanisms
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mechanisms[3]0.9the tagger's words
mechanismsmechanisms
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mechanisms[4]1.0the tagger's words
mechanismsmechanisms
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mechanisms[5]1.0the tagger's words
limitationsAwareness in monkeys could not be probed verbally, so the inference rests on the behavioural pattern alone.
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limitations[0]the tagger's words
limitationsNo subjective reports were collected from the monkeys; the subliminal thresholds were imported from the human procedure.
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limitations[1]the tagger's words
limitationsThe nonconscious effect may arise from low-level attentional orienting or residual visual traces rather than from a richer nonconscious process.
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limitations[2]the tagger's words
limitationsThe inference rests on a behavioural signature, not on any access to experience - the explanatory gap the paper cites is not closed by it.
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limitations[3]0.8the tagger's words
limitationsAwareness of the subliminal cue is established by self-report and a forced-choice test, both of which are null results rather than proof of absence.
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limitations[4]0.9the tagger's words
limitationsThe whole approach assumes the human dissociation transfers to another species; before this paper there was no evidence it does.
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limitations[5]1.0the tagger's words
transfersThey take the cross-over double-dissociation marker of nonconscious processing, developed and validated only in verbal human subjects, and transplant it into nonhuman-primate comparative research as a nonverbal test for awareness
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transfers[0]the tagger's words
transfersthe cross-over double dissociation of visual awareness, rebuilt as a nonverbal saccade task so that it can be run on a species that cannot be asked what it saw
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transfers[1]1.0the tagger's words
controlstwo NONPREDICTIVE cues, presented in both the supraliminal and the subliminal conditions
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eliminates: the mere presence of a cue as the explanation - if performance improved whenever any cue appeared, the effect would be arousal or alerting rather than the spatial information the cue carries · outcome: held
controls[0]0.95the tagger's words
conceptsdouble dissociation
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concepts[0]1.0the paper's words
conceptssingle dissociations
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concepts[1]1.0the paper's words
conceptssubliminal
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also in: subliminal
concepts[2]1.0the paper's words
conceptssupraliminal
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also in: supraliminal
concepts[3]1.0the paper's words
conceptsnonconscious processing
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concepts[4]0.85the paper's words
conceptsvisual awareness
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concepts[5]1.0the paper's words
conceptsspatial-cueing paradigm
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concepts[6]1.0the paper's words
conceptsexplanatory gap
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concepts[7]1.0the paper's words
conceptstheory of mind
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concepts[8]1.0the paper's words
sensesawareness - the reportable kind, defined operationally by what a subject says it saw and by a forced-choice test at chance
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senses[0]0.9the tagger's words
sensesawareness - the behavioural kind, defined by a crossing of two opposite performance signatures and requiring no report at all
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senses[1]1.0the tagger's words
sensesconsciousness - the demanding sense requiring language or theory of mind, which would confine it to humans and a few great apes
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senses[2]1.0the tagger's words
sensesconsciousness - the neuroanatomical sense, on which birds and octopuses may qualify
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senses[3]1.0the tagger's words
alternativesclaim_type: existence
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a second reading reaching the same value. Recorded: existence — note: a second reading reaching the same value from a different sentence. Not applied - a person decides.
alternatives.claim_typethe tagger's words
alternativesphenomenon: cross-over double dissociation between conscious and nonconscious visual processing
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a second reading DIFFERING from the recorded value. Recorded: double dissociation of visual awareness in nonhuman primates — note: a second reading, verbatim-verified, disagreeing with the value on the record. Not applied - a person decides.
alternatives.phenomenonthe tagger's words
claim_relationsextends → the single-dissociation literature on animal awareness (blindsight analogues, implicit/explicit memory, binocular rivalry)
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claim_relations[0]1.0the tagger's words
claim_relationscontradicts → the position that consciousness requires language or a human-like theory of mind
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claim_relations[1]0.8the tagger's words
claim_relationsbuilds-on → the spatial-cueing paradigm ( 37 - 40 )
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claim_relations[2]1.0the tagger's words
concept_edgesvisual awareness —operationalised-as→ double dissociation
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strength of the relation: 1 Audit 2026-09-28: DOUBTFUL - read against its full span and kept, but the double-dissociation framework is USED TO disentangle conscious from nonconscious processing; operationalised-as is loose.
concept_edges[0]the tagger's words
concept_edgessubliminal —contrasted-with→ supraliminal
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strength of the relation: 1
concept_edges[1]the tagger's words
concept_edgesvisual awareness —distinct-from→ nonconscious processing
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strength of the relation: 1 Audit 2026-09-28: DOUBTFUL - read against its full span and kept, but the span writes 'conscious' and 'nonconscious processing' / 'aware', not 'visual awareness'.
concept_edges[2]the tagger's words
concept_edgesnonconscious processing —operationalised-as→ subliminal
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strength of the relation: 1 Audit 2026-09-28: DOUBTFUL - read against its span and kept, but subliminal cueing is the method for nonconscious processing; operationalised-as is loose.
concept_edges[3]the tagger's words
content_tagslinear mixed model
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content_tags.statistics[0]the tagger's words
content_tagsF-test
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also in: F-test
content_tags.statistics[1]the tagger's words
content_tagscorrelation
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also in: correlation
content_tags.statistics[2]the tagger's words
content_tagsbinomial test
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also in: binomial test
content_tags.statistics[3]the tagger's words
content_tagschi-squared test
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content_tags.statistics[4]the tagger's words
content_tagschi-squared test
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content_tags.statistics[5]the tagger's words
content_tagsstandard error of the mean
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content_tags.statistics[6]the tagger's words
content_tagsmean and standard deviation
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content_tags.statistics[7]the tagger's words
content_tagslearning-slope correlation
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content_tags.statistics[8]the tagger's words
content_tagsanova
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also in: anova
content_tags.statistics[9]1.0the tagger's words
content_tagsstatistical-significance
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content_tags.statistics[10]1.0the tagger's words
content_tagsbinomial-test
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also in: binomial-test
content_tags.statistics[11]1.0the tagger's words
content_tagsSubliminal incongruent cues slowed human target detection relative to baseline even though participants reported and tested as unaware of them, establishing nonconscious processing.
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content_tags.findings[0]the tagger's words
content_tagsRhesus monkeys in the subliminal condition were significantly slower to detect the target than in the baseline with two nonpredictive subliminal cues.
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content_tags.findings[1]the tagger's words
content_tagsLike the human participants, the monkeys failed to ignore the subliminal cues and showed no signs of learning.
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content_tags.findings[2]the tagger's words
content_tagsSupraliminal-condition participants who reported no awareness of the cue's predictive value were not faster than baseline (324 ms against 312 ms).
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content_tags.findings[3]the tagger's words
content_tagsCue awareness, not cue saliency (the cue stayed hard to see at 33 ms), appears to be the critical factor in performance.
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content_tags.findings[4]the tagger's words
content_tagsPerformance stayed significantly below chance on trials where participants reported no awareness of the cue's location.
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content_tags.findings[5]the tagger's words
content_tagsRhesus monkeys show a double-dissociation pattern remarkably similar to that of humans.
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content_tags.findings[6]the tagger's words
content_tagsHumans are much faster with a visible incongruent cue than with an invisible one - the conscious condition helps.
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content_tags.findings[7]1.0the tagger's words
content_tagsThe same cue presented invisibly does the opposite: participants who saw nothing were nevertheless slowed relative to baseline.
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content_tags.findings[8]1.0the tagger's words
content_tagsThe double-dissociation signatures were first established in humans across seven experiments and then found in similar form in rhesus monkeys.
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content_tags.findings[9]0.9the tagger's words
content_tagsEarlier work established only single dissociations, which show animals have properties resembling conscious or nonconscious processing without separating the two.
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content_tags.findings[10]1.0the tagger's words
content_tagsspatial-cueing paradigm
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content_tags.definitions[2]the tagger's words
content_tagsforced-guessing paradigm
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content_tags.definitions[3]the tagger's words
content_tagscross-over double dissociation
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content_tags.definitions[6]1.0the tagger's words
content_tagssupraliminal stimuli
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content_tags.definitions[8]1.0the tagger's words
content_tagssubliminal stimuli
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content_tags.definitions[9]1.0the tagger's words
content_tagsresponse time
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also in: response time
content_tags.variables[2]the tagger's words
content_tagsgaze frequency
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content_tags.variables[4]the tagger's words
content_tagsmean = 43.5% correct, P = 0.00018 — Monkeys' below-chance accuracy in the subliminal forced-guessing condition and its combined binomial probability
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content_tags.quantities[0]the tagger's words
content_tags42% versus 25% — Share of human participants falling in the lowest binomial p-value quartile versus the share expected by chance
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content_tags.quantities[1]the tagger's words
content_tags94.6% correct, SD = 3.7% — Human accuracy in the supraliminal forced-guessing condition (eye-tracking modality)
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content_tags.quantities[2]the tagger's words
content_tags4 monkeys, 145 humans — Total subjects across the seven experiments
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content_tags.quantities[3]the tagger's words
content_tags75% — Participants who became aware of the cues and used them when only 20% of trials carried a supraliminal cue
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content_tags.quantities[4]the tagger's words
content_tags250 ms versus 17/33 ms — Cue exposure durations defining the supraliminal and subliminal conditions
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content_tags.quantities[5]the tagger's words
content_tags1.2% — Human error rate, which did not differ between supraliminal and subliminal conditions
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content_tags.quantities[6]the tagger's words
content_tags50% — Chance level in the forced-guessing task, above which performance requires using the incongruent cue
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content_tags.quantities[7]the tagger's words
content_tags250 ms vs 17/33 ms — supraliminal versus subliminal cue durations - the manipulation the whole design turns on
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content_tags.quantities[8]1.0the tagger's words
content_tagsseven experiments — size of the series, humans then monkeys
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content_tags.quantities[9]1.0the tagger's words
content_tags200 ms fixation — gaze hold required to register a response, the nonverbal substitute for a button press
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content_tags.quantities[10]1.0the tagger's words

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