Tonotopic organization of human auditory cortex — TEI-tagged view

Humphries, Liebenthal, Binder · 2010 · neuroscience · doi:10.1016/j.neuroimage.2010.01.046 · full text at the source · TEI xml · tagged edition · index · concepts · authors
Where this record's text came from. faceted 2026-08 by reading the locally held text (xml); every evidence string verified as an exact substring of that text before merging. Replication status is 'contested' rather than 'replicated' because the paper both claims confirmation of two mirror-symmetric gradients and states that the literature disagrees — 'Results and interpretations from these studies have varied considerably.' and 'The orientation of these two principal gradients is thus roughly perpendicular to HG, rather than parallel as previously assumed.' The gradient count is corroborated; the orientation is disputed. The paper also reports high between-subject variability ('The maps in our study did show a high degree of variability across subjects and across hemispheres.'), which I treated as a result rather than a limitations entry since it is presented as a finding supporting prior cytoarchitectonic work. Homology to macaque areas is stated as likely, not demonstrated, so I did not encode it as a claim relation to specific macaque papers. 2026-08-12 and 2026-08-14 schema-0.3 passes: every quote verified as an exact substring of the staged text and anchored at merge time; concept terms verified present; every edge endpoint an existing node.
Other fields define these differently the same word, defined another way in another discipline
fmri 2 fields
Tags on this paper every tag the record carries, whether or not the words occur in the text below — a number is how many papers in the corpus carry that tag; a tag with no number is unique to this paper
tonotopy2 cochleotopy frequency-selective response F test15 two principal gradients posterior lateral gradient area CL posterior parabelt macaque areas A1 and R core2 belt surface-based2 volume-based analysis cortical folding physical proximity retinotopic gradients receptive field gradients in auditory cortex planum temporale2 tuning bandwidth area CM cytoarchitectonic fmri24

Our tag layer for this paper

Cross-disciplinary analysis
content-tags & facets: phenomenon · claim-type · cross-domain-transfer · mechanism · claim-relation · replication-status · limitation — grounded in the full text
phenomenontonotopic organization of human auditory cortex— “The organization of tonotopic fields in human auditory cortex was investigated using functional magnetic resonance imaging.”
claim-typeexistence— “Two mirror-symmetric frequency gradients were found extending along an anterior-posterior axis from a zone on the lateral aspect of Heschl's gyrus (HG), which responds preferentially to lower frequencies, toward zones posterior and anterior to HG that are sensitive to higher frequencies.”
method — evidence— “FMRI data were collected from 8 right-handed subjects (7 male, 1 female; ages 20-37 years).”
object of study — evidence— “The organization of tonotopic fields in human auditory cortex was investigated using functional magnetic resonance imaging.”
research question — evidence— “In particular, we were interested in addressing several open questions that have not been completely resolved, such as the number of frequency gradients, the orientation of the gradients relative to HG, and the variability of the gradients across subjects.”
contribution — evidence— “The orientation of these two principal gradients is thus roughly perpendicular to HG, rather than parallel as previously assumed.”
transferfrom macaque single-unit electrophysiology: the core/belt field layout, used as the template against which a human fMRI map is read and labelled— “In the auditory cortex of nonhuman primates, each of the three divisions of primary “core” cortex – A1, R and RT – exhibit tonotopic gradients that are mirror symmetric to each other”transfers[0]
transferfrom visual neuroscience: retinotopy used as the contrast case for how a receptive-field gradient behaves at an areal boundary— “In contrast to the visual cortex, where retinotopic gradients reverse direction at the boundary between primary and secondary cortices, the receptive field gradients in auditory cortex run parallel to the primary/secondary (i.e., core/belt) boundary, such that the two mirror symmetric gradients observed in A1 and R extend laterally to belt…transfers[1]
mechanismcortical folding pushes the anterior and posterior high-frequency zones close together in 3-D, so volume-based smoothing and inter-subject variation blur them into one, hiding the perpendicular gradients from earlier studies— “The cortical folding that produces HG causes the high-frequency zones in the sulci posterior and anterior to HG to be physically very close in 3-dimensional space. This physical proximity, together with inter-subject variation in the exact location of the sulci and spatial smoothing used for generating group maps, may have caused these…mechanisms[0]
mechanismflattening the superior temporal plane onto a surface model pulls the buried sulcal regions apart, which is what makes the perpendicular topology visible— “Flattening the superior temporal plane with surface modeling separates these regions and allows a more accurate analysis of topological relations running perpendicular to the gyrus.”mechanisms[1]
mechanismcortical folding puts the anterior and posterior high-frequency zones physically close together, so volume-based analysis and smoothing blur them into one— “The cortical folding that produces HG causes the high-frequency zones in the sulci posterior and anterior to HG to be physically very close in 3-dimensional space. This physical proximity, together with inter-subject variation in the exact location of the sulci and spatial smoothing used for generating group maps, may have caused these…mechanisms[2]
mechanismflattening the superior temporal plane separates regions that are adjacent in 3-D, making gradients perpendicular to the gyrus visible— “Flattening the superior temporal plane with surface modeling separates these regions and allows a more accurate analysis of topological relations running perpendicular to the gyrus.”mechanisms[3]
mechanismsparse sampling: the scanner runs only in a 1.6 s silent gap after the 8 s tone sequence, so scanner noise never overlaps the stimulus— “A trial lasted 9.6 seconds and started with an 8-second tone sequence followed by a 1.6-second gap. MRI scanning was performed only during the 1.6-second gap to prevent interference from scanner noise during presentation of the stimulus.”mechanisms[4]
mechanismstochastic alternation between two- and three-tone bursts holds the cortical response up, since auditory neurons adapt after onset— “This alternating pattern was designed to provoke large responses in auditory cortical neurons, which tend to have increased firing rates at stimulus onset but often return to baseline before the end of the stimulus”mechanisms[5]
mechanismtone ratios chosen so the composite produces no harmonic relationship and no missing-fundamental pitch— “These ratios were selected so that the composite tones would have no harmonic relationships and would not produce a “missing fundamental” pitch, and so that the five tones comprising the stimulus train would be approximately equidistant in pitch space.”mechanisms[6]
mechanismbest frequency is interpolated between the six tested bands by weighting adjacent conditions, giving finer resolution than the design hasmechanisms[7]
mechanismlandmark-based surface alignment rather than volumetric alignment, for spatial precision in the temporal lobe
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mechanisms[8]
mechanismnon-parametric permutation testing because eight subjects will not support a parametric group test
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mechanisms[9]
mechanismseveral sessions per subject, to buy single-subject statistical power and a test-retest estimate at the same time
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mechanisms[10]
replicates Formisano et al. (2003); Talavage et al. (2004); Woods et al. (2009)
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claim_relations[0]
contradicts Formisano et al. (2003), on the orientation of the gradients relative to HG
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claim_relations[1]
contradicts Woods et al. (2009), on labelling the posterior lateral gradient as parabelt
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claim_relations[2]
partially-supports Formisano et al. 2003: two mirror-symmetric gradients, but running along the axis of HG
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claim_relations[3]
refines Talavage et al. 2004: three gradients described as narrow lines between high and low poles
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claim_relations[4]
supports Woods et al. 2009: one low-frequency area on HG flanked by two high-frequency areas
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claim_relations[5]
contradicts Woods et al. 2009: the posterior lateral field is parabelt
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claim_relations[6]
contradicts the early two-frequency studies: a single gradient running along the axis of HG
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claim_relations[7]
qualifies their own reading that these central regions are more sharply tuned
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claim_relations[8]
supports macaque evidence that area CM prefers higher than average frequencies without a gradient
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claim_relations[9]
sensetonotopy - a gradient of preferred frequency mirroring the cochlea; explicitly a map of one stimulus dimension, not of any other response property
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senses[0]
sensegradient - as a field bounded by high and low bands, not as a line between two poles; the paper says this reading is itself a source of the literature's disagreement
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senses[1]
sensefrequency-selective - defined operationally as a significant F test across the six tested frequencies, carrying no claim about tuning shape
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senses[2]
sensetuning bandwidth - here a ratio of preferred to non-preferred BOLD amplitude, which the paper scare-quotes and then declines to read as tuning width
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senses[3]
sensecore and belt - cytoarchitectonic and connectional divisions, explicitly not divisions defined by the frequency map being measured
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senses[4]
sensecorresponds to / homologous to - used hedged throughout for human-macaque field matches, never as identity
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senses[5]
controlthe intensity of every tone sequence normalised to A-weighted 76.3 dB — held — rules out a loudness difference between sequences - otherwise a tonotopic response could be a response to how loud each frequency happened to be
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controls[0]
differs inFormisano et al. (2003), on the orientation of the gradients relative to HG — the ORIENTATION of the gradients relative to HG: THESE GRADIENTS ARE ORIENTED APPROXIMATELY ALONG THE AXIS OF HG, WHEREAS THE GRADIENTS WE OBSERVED WERE PERPENDICULAR TO THE LONG AXIS OF HG
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claim_relations[1]
differs inWoods et al. (2009), on labelling the posterior lateral gradient as parabelt — where the parabelt is and whether it is tonotopic at all: THERE IS LITTLE EVIDENCE FOR TONOTOPIC ORGANIZATION IN THE PARABELT OF MONKEYS, and RECENT HUMAN CYTOARCHITECTONIC DATA SUGGEST THAT THE PARABELT IS LOCATED MORE VENTRALLY IN THE UPPER BANK OF THE SUPERIOR TEMPORAL SULCUS
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claim_relations[2]
differs inWoods et al. 2009: the posterior lateral field is parabelt — which area the same patch of cortex IS - the two papers show A SIMILAR DIAGRAM and label it differently: AREA CL here, POSTERIOR PARABELT there, and the criterion offered is that IN THE MONKEY, THIS GRADIENT DOES NOT EXTEND INTO ADJACENT AREAS; THEREFORE, ITS BOUNDARIES ARE EASILY DETERMINED
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claim_relations[6]
differs inthe early two-frequency studies: a single gradient running along the axis of HG — the analysis method, named as decisive across the whole prior literature: ONE THAT WE BELIEVE IS CRITICAL IS THAT THE STUDIES REPORTING GRADIENTS PERPENDICULAR TO HG ALL USED SURFACE-BASED RATHER THAN VOLUME-BASED ANALYSIS
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claim_relations[7]
differs intheir own reading that these central regions are more sharply tuned — which side of the difference moves: the sharper tuning is DRIVEN PRIMARILY BY INCREASES IN RESPONSE TO THE PREFERRED FREQUENCY RATHER THAN DECREASES IN THE RESPONSE TO THE NON-PREFERRED FREQUENCIES, so the pattern IS CORRELATED WITH THE OVERALL STRENGTH OF THE RESPONSE and need not be sharper tuning
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claim_relations[8]
notetonotopy —same-as→ cochleotopy — given as alternate names in one clause, unhedged - a positive control for the join key
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concept_edges[0]
edgefrequency-selective response —operationalised-as→ F test
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concept_edges[1]
differs insurface-based —contrasted-with→ volume-based analysis — THE AXIS IS THE ANALYSIS METHOD, AND IT IS OFFERED AS THE EXPLANATION OF THE DISAGREEMENT. 'Although there are many differences in methodology between these studies, one that we believe is critical is that the studies reporting gradients perpendicular to HG all used surface-based rather than volume-based analysis.' The authors name the one difference they think decides the result out of many they could have named - which is the analyst decision this project exists to make legible.
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concept_edges[2]
notesurface-based —contrasted-with→ volume-based analysis — the contrast the paper says explains the whole literature's disagreement
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concept_edges[2]
effect directioncortical folding —causes→ physical proximity — increases
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concept_edges[3]
weightcortical folding —causes→ physical proximity — 0.95
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concept_edges[3]
auditposterior lateral gradient —same-as→ area CL — Audit 2026-09-28: DOUBTFUL - read against its full span and kept, but the gradient 'MOST LIKELY corresponds to' area CL; same-as is hedged.
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concept_edges[5]
noteposterior lateral gradient —same-as→ area CL — most likely corresponds to
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concept_edges[5]
weightposterior lateral gradient —same-as→ area CL — 0.65
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concept_edges[5]
noteposterior lateral gradient —distinct-from→ posterior parabelt — a labelling dispute with Woods et al., settled by appeal to a different evidence type
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concept_edges[6]
weightposterior lateral gradient —distinct-from→ posterior parabelt — 0.85
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concept_edges[6]
differs incore —distinct-from→ belt — THE AXIS IS WHAT DEFINES THE BOUNDARY. 'with the boundary between core and belt defined by cytoarchitectonic and connectivity patterns rather than tonotopic gradients.' The two regions are not distinguished by the property the paper measures - they are distinguished by a different property entirely, so a tonotopic result cannot confirm or refute the division it is described against.
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Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
concept_edges[7]
notecore —distinct-from→ belt — the boundary is stipulated from cytoarchitecture and connectivity, not from the phenomenon being mapped
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concept_edges[7]
noteretinotopic gradients —contrasted-with→ receptive field gradients in auditory cortex — a structural contrast between sensory systems sharing almost no vocabulary
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concept_edges[8]
outcome (stated by the tagger, no quotation)control — heldcontrols[0]
for which uselimitation — the core/belt boundary drawn on the map: THE EXACT BOUNDARY BETWEEN CORE AND BELT CANNOT BE DETERMINED BASED SOLELY ON FREQUENCY RESPONSE, THUS THIS LINE IS AN ESTIMATE GUIDED BY THE KNOWN TOPOGRAPHICAL ORGANIZATION IN THE MACAQUE
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limitations[0]
for which uselimitation — the claimed absence of an RT-like gradient: THE LACK OF FREQUENCY-SELECTIVE ACTIVATION IN THIS REGION COULD BE DUE TO LOW SENSITIVITY TO THE STIMULI USED IN THE CURRENT STUDY
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limitations[1]
for which uselimitation — the group significance test, and the paper says what it did about it: BECAUSE OF THE SMALL NUMBER OF SUBJECTS, SIGNIFICANCE WAS CALCULATED USING A NON-PARAMETRIC PERMUTATION TEST
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limitations[2]
for which uselimitation — tuning estimates at the edges of the tested range: POINTS WERE ALSO EXCLUDED IF THEIR LARGEST RESPONSE WAS TO THE LOWEST (200 Hz) OR HIGHEST (6400 Hz) FREQUENCY
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limitations[3]
for which uselimitation — the same scope as limitations[1], left open: ALTERNATIVELY, IT IS POSSIBLE THAT AREA RT DOES NOT EXIST IN HUMANS - a sensitivity failure and a real absence are not distinguished
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limitations[4]
for which uselimitation — the sharper-tuning reading: THESE PATTERNS GENERALLY APPEAR TO BE CORRELATED WITH THE OVERALL STRENGTH OF THE RESPONSE
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limitations[5]
for which uselimitation — the same scope as limitations[0] - the core/belt line, imported from macaque and cytoarchitecture rather than measured here
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limitations[6]
for which uselimitation — the parietal operculum activation, which is not a finding: THIS ACTIVATION IS PRIMARILY AN ARTIFACT OF THE MAPPING BETWEEN VOLUME AND SURFACE SPACES DUE TO PARTIAL VOLUME EFFECTS
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limitations[7]
for which uselimitation — any claim about where the border between gradients falls: THE LOCATION OF THE LOW-FREQUENCY BORDER BETWEEN THE TWO GRADIENTS VARIED IN RELATION TO HG, with A HIGH DEGREE OF VARIABILITY ACROSS SUBJECTS AND ACROSS HEMISPHERES
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limitations[8]
for which uselimitation — voxel counts and preferred frequency across sessions: SOME INTER-SESSION VARIABILITY WAS SEEN IN THE NUMBER OF SIGNIFICANTLY ACTIVATED VOXELS AND THEIR PREFERRED FREQUENCY
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limitations[9]
for which uselimitation — the population every map describes: FMRI DATA WERE COLLECTED FROM 8 RIGHT-HANDED SUBJECTS (7 MALE, 1 FEMALE; AGES 20-37 YEARS)
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limitations[10]
for which uselimitation — the attentional state the tonotopic maps were measured in: SUBJECTS WERE INSTRUCTED TO LIE STILL AND LISTEN TO THE TONES. NO SPECIFIC TASK WAS PERFORMED DURING THE EXPERIMENT
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limitations[11]
duplicate ofmechanism — mechanisms[3]
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mechanisms[1]
duplicate ofmechanism — mechanisms[0]
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mechanisms[2]
replicationcontested
limitationThe core/belt boundary drawn on the homology map cannot be established from frequency responses alone and is an estimate imported from macaque anatomy and human cytoarchitecture.
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limitations[0]
limitationThe absence of a gradient corresponding to macaque area RT may be a sensitivity failure of the stimuli rather than evidence that the area is absent in humans.
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limitations[1]
limitationThe sample was small enough to force non-parametric permutation testing for the group analysis.
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limitations[2]
limitationthe tested range truncates the tuning estimate, so points preferring 200 or 6400 Hz were excluded from that analysis
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limitations[3]
limitationan absence the authors decline to resolve: no anterior RT-like gradient, which may be the stimuli or may be the species
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limitations[4]
limitationthe sharper-tuning reading is undercut by the authors themselves, because the effect tracks overall response strength
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limitations[5]
limitationthe core/belt boundary cannot be fixed from frequency response alone, so the line drawn is an estimate imported from other evidence
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limitations[6]
limitationapparent parietal operculum activation is an artifact of volume-to-surface mapping, not a finding
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limitations[7]
limitationhigh variability across subjects and hemispheres in where the low-frequency border between gradients falls
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limitations[8]
limitationinter-session variability attributed partly to head movement and partly to unmeasured attention and alertness
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limitations[9]
limitationeight subjects, seven of them male
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limitations[10]
limitationno task was performed, so attentional state during scanning is uncontrolled by design
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limitations[11]
Content tags
content-observations: finding · quantity · variable · definition · key-term — grounded in the full text
findingTwo mirror-symmetric frequency gradients run anterior-posterior from a low-frequency zone on lateral Heschl's gyrus out to high-frequency zones in front of and behind it.
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content_tags.findings[0]
findingThese principal gradients lie perpendicular to HG, overturning the prevailing assumption that they run parallel to it.
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content_tags.findings[1]
findingA third, smaller gradient was found on the lateral posterior superior temporal gyrus.
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
content_tags.findings[2]
findingThe human maps are argued to be closely homologous to nonhuman primate auditory cortex, with the two main gradients corresponding to macaque A1 and R.
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
content_tags.findings[3]
findingIndividual tonotopic maps were highly reproducible across separate scanning sessions.
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
content_tags.findings[4]
findingInspection of unnormalized data suggests the regional differences are driven mainly by a larger response to the preferred frequency rather than a smaller response to non-preferred ones.
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
content_tags.findings[5]
evidence status notefinding — QUOTE RECOVERED FROM THE PAPER'S OWN CHARACTERS. The recorded quote was LETTERS-ONLY IDENTICAL to this stretch of the staged text but not an exact substring of it - spaces, hyphens, ligature glyphs or mathematical operators differed, so it could never anchor. This is the file's own text between the same two letters: not produced by a rule about the damage, copied out of the file. Round-trip checked (the recovered span is letters-only equal to the original) and length-checked. The text as originally recorded is kept in evidence as extracted.content_tags.findings[5]
findingtwo mirror-symmetric gradients run roughly perpendicular to HG, not parallel as previously assumed
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
content_tags.findings[6]
findinga low-frequency zone on lateral HG is flanked anteriorly and posteriorly by high-frequency zones in every subject
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
content_tags.findings[7]
findinga third, smaller gradient runs along lateral posterior STG, mainly in the left hemisphere
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
content_tags.findings[8]
findingfrequency-selective responses were confined to the superior temporal plane in both hemispheres
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
content_tags.findings[9]
findingmaps are highly reproducible across sessions within a subject
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
content_tags.findings[10]
findingthe unnormalized data suggest the difference is driven mainly by a larger preferred response rather than a smaller non-preferred one
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
content_tags.findings[11]
findingno distinct frequency gradient in area CM, agreeing with macaque evidence for an overall high-frequency preference there
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
content_tags.findings[12]
findingprior imaging results and their interpretations varied considerably
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
content_tags.findings[13]
findingfour previous studies reached four different conclusions, one of them denying the gradients are gradients at all
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
content_tags.findings[14]
quantity8 — number of subjects scanned, with sex split and age range
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
content_tags.quantities[0]
quantity392 trials total — trials per scanning session, from 7 runs of 56 trials
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
content_tags.quantities[1]
quantity.834 (SD = 0.083) — mean correlation of a subject's tonotopy map between scanning sessions, i.e. test-retest stability
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
content_tags.quantities[2]
quantity20 to 30 mm 3 — range of the per-subject cluster-size threshold used for multiple-comparison correction
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
content_tags.quantities[3]
evidence status notequantity — QUOTE RECOVERED FROM THE PAPER'S OWN CHARACTERS. The recorded quote was LETTERS-ONLY IDENTICAL to this stretch of the staged text but not an exact substring of it - spaces, hyphens, ligature glyphs or mathematical operators differed, so it could never anchor. This is the file's own text between the same two letters: not produced by a rule about the damage, copied out of the file. Round-trip checked (the recovered span is letters-only equal to the original) and length-checked. The text as originally recorded is kept in evidence as extracted.content_tags.quantities[3]
quantity76.3 dB — A-weighted intensity to which every tone sequence was normalised
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
content_tags.quantities[4]
quantity9.6 seconds — trial length: an 8-second sound followed by a 1.6-second silent gap in which the scan was acquired
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
content_tags.quantities[5]
quantity200, 400, 800, 1600, 3200, 6400 Hz — the six centre frequencies tested
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
content_tags.quantities[6]
quantity8 right-handed subjects, 7 male and 1 female, ages 20-37 — the sample
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
content_tags.quantities[7]
quantity.834, SD 0.083 — mean between-session correlation of the tonotopy maps
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
content_tags.quantities[8]
quantitybetter than 20 dB HL from 250 to 8000 Hz — audiometric inclusion criterion
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
content_tags.quantities[9]
quantity20 to 30 mm3 — cluster threshold, which varied by subject because smoothness was estimated per subject
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
content_tags.quantities[10]
quantity8 s stimulus then a 1.6 s scanning gap, 9.6 s per trial — the sparse-sampling timing
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
content_tags.quantities[11]
quantity3000 Hz — Formisano's highest frequency, offered as the reason that study may have missed medial anterior high-frequency cortex
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
content_tags.quantities[12]
quantity3600 Hz, 4 Hz presentation rate — Woods' ceiling and rate, against 6400 Hz and 10 Hz here
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
content_tags.quantities[13]
variablefrequency of maximum response (dependent)
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
content_tags.variables[1]
variablemean BOLD signal (dependent)
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
content_tags.variables[2]
definitiontuning bandwidth
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
content_tags.definitions[1]
definitionfrequency-selective response
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
content_tags.definitions[2]
definitiontonotopy
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
content_tags.definitions[3]
definitionfrequency-selective response
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
content_tags.definitions[4]
key-termstonotopy, cochleotopy, Heschl, planum temporale, superior temporal gyrus, mirror-symmetric, core, belt, area CL, area CM, posterior parabelt, macaque areas A1 and R, frequency-selective response, tuning bandwidth, surface-based, volume-based analysis, cortical folding, retinotopic gradients, cytoarchitectonic, permutation test
Concepts
terms this paper carries, each with the sentence it was cut from — the layer that joins one paper to another. Terms with no quote yet are on CONCEPTS.html.
conceptcortical folding — not defined in this paper's own words; the gyral geometry that creates the confound
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
concepts[13]
conceptphysical proximity — not defined in this paper's own words; the closeness in 3-D of the two high-frequency zones flanking HG
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
concepts[14]
Statistics — cross-field join keys
field-invariant statistical terms, used in their statistical sense (gated on a number / symbol / test). A p-value or a standard deviation means the SAME thing in physics, biology, psychology and sociology — so these are the strongest cross-field join keys. Grounded verbatim.
general linear model as written: A general linear model
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content_tags.statistics[0]
F-test F as written: F-tests
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
content_tags.statistics[1]
multiple comparison correction p as written: cluster thresholded to correct for multiple comparisons at an alpha level of .05
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
content_tags.statistics[2]
ANOVA as written: one-way ANOVA
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
content_tags.statistics[3]
permutation test as written: non-parametric permutation test
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
content_tags.statistics[4]
evidence status notepermutation test — QUOTE RECOVERED FROM THE PAPER'S OWN CHARACTERS. The recorded quote was LETTERS-ONLY IDENTICAL to this stretch of the staged text but not an exact substring of it - spaces, hyphens, ligature glyphs or mathematical operators differed, so it could never anchor. This is the file's own text between the same two letters: not produced by a rule about the damage, copied out of the file. Round-trip checked (the recovered span is letters-only equal to the original) and length-checked. The text as originally recorded is kept in evidence as extracted.content_tags.statistics[4]
correlation as written: mean between-session correlation coefficient
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
content_tags.statistics[5]
F test across six frequency conditions
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
content_tags.statistics[6]
cluster thresholding
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
content_tags.statistics[7]
non-parametric permutation test
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
content_tags.statistics[8]
between-session correlation coefficient
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
content_tags.statistics[9]
Second readings
a facet read a second time, independently, from its own sentence — and whether the two readings agree
methodfMRI with sparse sampling, six multi-tone frequency bands from 200 to 6400 Hz, analysed on flattened cortical surfaces with landmark-based alignment — a second reading DIFFERING from the recorded value; recorded: fmri
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
alternatives.method
phenomenontonotopy - a cortical gradient of preferred frequency mirroring the cochlea — a second reading DIFFERING from the recorded value; recorded: tonotopic organization of human auditory cortex
Quotation withheld — this paper is not ours to reproduce, and the page has already quoted as much of it as the rights layer allows. The tag, its field and its confidence stand; the sentence is at the publisher.
alternatives.phenomenon

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